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Nathan Hughes available for England's showdown with Australia

Nathan Hughes of England takes on Faf de Klerk of South Africa at Twickenham
Image: Nathan Hughes is set for his first start for England

Nathan Hughes is available for England's final autumn international against Australia on Saturday after taking a full part in training.

The Wasps star missed the 27-14 victory over Argentina with a foot injury but has been cleared to replace Billy Vunipola in the back row against the Wallabies at Twickenham in what would be his full Test debut.

Vunipola has been ruled out for at least three months after undergoing knee surgery, jeopardising his participation in the Six Nations, and Eddie Jones has already described the Fijian-born Hughes as the "obvious" replacement at No 8.

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Will Greenwood has played down comments made by Eddie Jones and Michael Cheika

"Nathan has done really well in camp since he joined, last week's injury aside," forwards coach Steve Borthwick said.

"Billy is a tremendous player who has done really well for us and has been a big figure in our team.

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"But injuries are an opportunity for other players and what we've seen throughout this series is players coming in and taking chances presented by injury.

"This is another one of those opportunities. We have a lot of quality back row forwards in the squad."

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Australia head coach Michael Cheika has accused his England counterpart Eddie Jones of 'vitriol'

Jack Clifford offers No 8 cover but having just made his comeback from ankle surgery, he must now observe the return to play protocols for concussion after taking a blow to the head for Harlequins against Bath on Sunday.

England defeated Argentina despite seeing wing Elliot Daly sent off in the fifth minute and for all the Pumas' inability to exploit their numerical advantage, it was a remarkable performance.

COVENTRY, ENGLAND - SEPTEMBER 04:  Nathan Hughes of Wasps breaks away with the ball during the Aviva Premiership match between Wasps and Exeter Chiefs at t
Image: Hughes has been outstanding for Wasps this season

"It was colossal. We had 14 men for 75 minutes but we still won a Test match against a team that's as strong as Argentina," defence coach Paul Gustard said.

"Everyone was raving about Argentina in the World Cup 12 months ago and they had a strong Rugby Championship.

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Eddie Jones is full of praise after his England side beat Argentina after going down to 14-men

"For 15 minutes we had 13 men. At the start of the second half, one bad defensive error led to a try and that's Test match rugby - you get punished for respects.

"We all felt unbelievably privileged to be part of a group that responded so courageously and heroically for a large portion of the game."

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